Trees are not just natural resources; they are also cultural ones that present unique challenges and opportunities for public historians. Trees can serve as important objects of memory, recalling past triumphs or tragedies. They can be the last living witness to important events or community stories. Trees can also be objects of preservation, sometimes as individuals, other times as stands or even forests, all of which can take on historical significance for people, sites, and institutions. But as living entities they defy the kind of permanent legal preservation applicable to buildings and other non-living historical objects. Furthermore, their organic fragility can actually make them significant problems for historical sites and local preservation activities. For example, communities have had to cope with extensive tree loss from storm and fire damage, and dying trees can drop limbs or topple over, creating considerable danger to people and resources. Climate-change-driven increasing storm intensity has also highlighted the ways that trees—however historical or beloved—can become considerable threats. The fourteen new, previously unpublished essays in this volume explore the many ways that trees are an integral part of public history practice and sites. The authors draw on a range of approaches and historiographies to look at how memories of race-based hate, patriotic stories, community identities, and changed places all have centered on trees. In addition to contributions from the volume editors, this collection features scholarship by Sonja DÜmpelmann Andrew Hurley, Carolyn M. Barske Crawford, Brian Dempsey, Liz Sargent, Sasha Coles, Mariaelena DiBenigno, Evan Haefeli, Krista McCracken, Alena Pirok, Christian Kosmas Mayer Alaina Scapicchio, and David Glassberg.